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Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Synonyms:

Rhizocarpon richardii (Lamy ex. Nyl.) Zahlbr. subsp. constrictum (Malme) Cl. Roux

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr. subsp. richardii

Rhizocarpon atlanticum I.M. Lamb.

Rhizocarpon constrictum Malme



Family: RHIZOCARPACEAE

Habitat: Xeric supralittoral  zone

Description

    Thallus crustose, potentially forming large mosaics, tightly encrusted on the rock surface, usually more or less circular, to 10 cm diam., areolate, cracked, network-forming, pale grey, grey, grey-brownish with grey-purple or grey-violet tinges, prothallus black, often large and well developed, markedly visible at the rim and between the areoles. Apothecia very numerous, 0.5-1 mm diam., inserted in the thallus, flat to slightly convex, surface uneven,, exciple thin, slightly paler, indistinct, black. Ascopores variable in shape and aspect (including from ascii and apothecia belonging to the same thallus), sometimes 1-septate  but often apparently submuriform with an incomplete secondary network suggesting muriform septa, but seldom truly muriform, colourless though some well mature specimen may become pale grey-brown, 22-27 (32) x 11-14 µm according to literature, 22-27 x 10-15 µm according to our measures. Photobiont: Chlorococcoid. C+ red (medulla), KC+ red (medulla), K+/- yellow, Pd+/- orange (Reactions vary depending on the gyrophoric, stictic and norstictic acid contents). Medulla: I+ indigo. Rather common on siliceous rocks, notably smooth ones, or on large pebbles on the foreshore where its aspect of irregular tiles, alternating dark and pale zones, often with purple tinge is characteristic of this species.

_ Not to be confused with Rhizocarpon reductum Th. Fr. that is C-, paler, smaller, less draughtboard-shaped, more common, with muriform ascospores and not especially maritime.

_ Not to be confused with taxa belonging to Buellia aethalea s.l. sharing the same type of well developed black prothallus but K+ yellow > red and with ascospores strictly 1-septate. On maritime rocks, see also Buellia stellulata  that is C-, K+ yellow and Pd+ yellow and much smaller (with 1-septate ascospores as in all Buellia species). 

_ It has been described Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr. subsp. richardii which would have spores with a gray-brown submural appearance when mature and which above all would not be maritime. Currently this subsp. is not taken into account by the current authors.

 





Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Crozon
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Asque Crozon 7
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Camaret
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Cap de la Chèvre
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Pointe de Pen Hir 2
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Pointe de Pen Hir 1
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Ouessant 1
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Ouessant 2
Bernard BOUFFINIER

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Pointe de Dinan 1
Alain GERAULT

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Pointe de Dinan 2
Alain GERAULT

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Crozon 2
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Crozon 3
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Réactions au C Crozon 4
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Asque avec spores immatures Crozon 5
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Spores Crozon 6
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Spores Crozon 8
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Spores Crozon 9
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Spores Crozon 10
Jacques HAINE

Rhizocarpon richardii (Lamy ex Nyl.) Zahlbr.

Rhizocarpon richardii Pointe de Pen-Hir 3
Michel DAVID



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